Years after high school, Christine, Crystal, Alice, and Lisa return to the Finger Lakes to perform again as Chrysalis, their schooldays quartet. They discover that what still binds them together is not only their music but having to deal with their problematic mothers. One wonders what her deceased mother was like as a person, not just the "mother figure" she knew. Another thinks her mother lives her life recklessly. The parent perceived as controlling is also struggling to hide her Parkinson's from the outside world, and the fourth has burgeoning dementia. While the adult daughters are rediscovering harmony in their singing and in their friendships, the mothers form bonds of their own--bonds made of secrets and new discoveries--and ultimately they find answers that bring the mothers and their daughters to a deeper understanding of themselves and their relationships to each other.